Two clones of Chara cf. globularis Thuill. with orthotropic thalli were found in mosses protonemas and gametophytes cultures under air conditions and 100% soil moisture capacity. The orthotropic ascending filaments formation of Charales representatives in terrestrial envinronment was not registered before. This fact may be interesting in consideration with land plants origin clarification and their relationships with streptophyte algae.

Keywords: terrestrial algae, soil cultures, Chara, orthotropic growth

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